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When food runs short, they slowly become clustered together and enter their "gregarious" phase, culminating in an aggressive swarm. Prior to swarming, the locusts undergo a series of dramatic physical changes - their body colour darkens and their muscles grow stronger. ... They found that locusts behaving the most gregariously (in swarm-mode) had approximately three times more serotonin in their systems than their calm, solitary...
 locust swarm serotonin depression
Swarming locusts not only look different and act differently to solitary locusts, they also have much larger brains.


 locust brain swell collaboration social neuroscience insect swarm